Effects of Rice Husk Biochar and Corn Straw Biochar on Growth，Quality and Soil Nutrients of Huai Chrysanthemum

doi:10.15933/j.cnki.1004-3268.2026.09.003

Abstract

Abstract: To investigate the effects of rice husk biochar and corn straw biochar on the growth，quality，and soil nutrients of Huai chrysanthemum，three levels of rice husk biochar（150，300，450 kg/ha，numbered D1，D2，D3 in sequence）and three levels of corn straw biochar（300，450，600 kg/ha，numbered Y1，Y2，Y3 in sequence） were set up，with no application of rice husk biochar and corn straw biochar as a control（CK）.A total of seven treatments were conducted to study the effects of the two biochars on the agronomic traits，photosynthetic parameters，yield，medicinal components，soil available nutrients，and key enzyme activities of Huai chrysanthemum. The results showed that compared with CK，the application of two types of biochar significantly improved the plant height，maximum photochemical efficiency（Fv/Fm），stomatal conductance（Gs），transpiration rate（Tr），and total yield of Huai chrysanthemum. Among them，D3 and Y3 treatments showed a higher increase in plant height，with an increase of 21.2% and 22.5% respectively compared to CK；the stem thickness of Y2 treatment was the largest，increasing by 13.49% compared to CK；the optimal number of branches was in D2，with an increase of 56.23%；Y3 showed the highest increase in net photosynthetic rate（Pn），Gs，and Tr compared to CK，with increases of 116.91%，307.21%，and 210.76%，respectively.The yield performance was better in D2，Y2，and Y3 treatments，with significant increases of 57.39%，53.19%，and 59.13% compared to CK，respectively. In terms of quality，the total flavonoids of all biochar treatments were significantly increased by 18.05%—55.86% compared to CK.There was no significant difference in chlorogenic acid and 3，5‑O‑dicaffeoylquinic acid between rice husk biochar group and CK，while the content of hesperidin was significantly reduced by 17.42%—23.87%.The chlorogenic acid and 3，5‑O‑dicaffeoylquinic acid treated with corn straw biochar increased by 9.27%—22.06% and 18.78%—28.45% respectively compared to CK. The content of hesperidin significantly increased with increasing application rate，with Y3 treatment showing a significant increase of 16.87% compared to CK.In terms of soil indicators，both types of biochar can significantly increase the content of soil alkaline hydrolyzable nitrogen，available phosphorus，available potassium，as well as urease and sucrase activities overall.Among them，the content of alkaline hydrolyzable nitrogen，available phosphorus，and available potassium in soil treated with D3 and Y3 increased by 5.05%，9.06%，6.67% and 4.28%，6.68%，7.44% respectively compared to CK；the urease activity significantly increased by 228.41% and 321.59% compared to CK，respectively.The peak of sucrase activity appeared in D2 and Y2 treatments，which increased by 46.71% and 58.48% respectively compared to CK. In summary，D2，Y2，and Y3 treatments showed the pretty overall performance.The recommended amount of biochar for field cultivation of Huai chrysanthemum is 300 kg/ha of rice husk biochar or 450—600 kg/ha of corn straw biochar.

Key words: Huai chrysanthemum, Rice husk biochar, Corn straw biochar, Growth, Quality, Soil nutrients

摘要： 为探明稻壳炭与玉米秸秆炭对怀菊生长、品质及土壤养分的影响，以怀菊为研究对象，设置3个稻壳炭水平（150、300、450 kg/hm^²，依次编号D1、D2、D3），3个玉米秸秆炭水平（300、450、600 kg/hm^²，依次编号Y1、Y2、Y3），以不施稻壳炭与玉米秸秆炭为对照（CK），共7个处理开展田间试验，研究2种生物炭对怀菊农艺性状、光合参数、产量、药效成分及土壤速效养分、关键酶活性的影响。结果表明，相较于CK，施用2种生物炭均可显著提升怀菊株高、最大光化学效率（Fv/Fm）、气孔导度（Gs）、蒸腾速率（Tr）与总产量。其中D3、Y3 处理株高较高，分别较CK 提升21.2%、22.5%；Y2 处理茎粗最大，较CK 提高13.49%；D2处理分枝数最优，增幅达56.23%；Y3处理净光合速率（Pn）、Gs、Tr提升幅度最高，依次较CK提高116.91%、307.21%、210.76%。产量表现以D2、Y2、Y3 处理较优，较CK 分别显著增产57.39%、53.19%、59.13%。品质方面，各生物炭处理总黄酮含量较CK显著提高18.05%~55.86%。稻壳炭组绿原酸、3，5-O-二咖啡酰基奎宁酸含量与CK无显著差异，木犀草苷含量显著降低17.42%~23.87%；玉米秸秆炭处理的绿原酸、3，5-O-二咖啡酰基奎宁酸含量分别较CK提高9.27%~22.06%、18.78%~28.45%，木犀草苷含量随施用量递增显著上升，其中Y3处理较CK显著提升16.87%。土壤指标方面，2种生物炭整体上均能显著提升土壤碱解氮、速效磷、速效钾含量及脲酶、蔗糖酶活性。其中，D3与Y3处理土壤碱解氮、速效磷、速效钾含量分别较CK提升5.05%、9.06%、6.67%与4.28%、6.68%、7.44%，脲酶活性分别较CK显著提高228.41%、321.59%；蔗糖酶活性峰值出现在D2、Y2处理，较CK分别提升46.71%、58.48%。综上，D2、Y2、Y3处理综合表现较优，怀菊大田栽培生物炭建议用量为稻壳炭 300 kg/hm^²或玉米秸秆炭450~600 kg/hm^²。

关键词: 怀菊, 稻壳炭, 玉米秸秆炭, 生长, 品质, 土壤养分

CLC Number:

S567.7

Fu Shenglan, Gu Caihua, Zhang Yanlin, Sun Wanhui, Xin Longfei. Effects of Rice Husk Biochar and Corn Straw Biochar on Growth，Quality and Soil Nutrients of Huai Chrysanthemum[J]. Journal of Henan Agricultural Sciences, 2026, 55(9): 26-34.

扶胜兰, 谷彩花, 张艳玲, 孙万慧, 信龙飞. 稻壳炭与玉米秸秆炭对怀菊生长、品质及土壤养分的影响[J]. 河南农业科学, 2026, 55(9): 26-34.

Figures/Tables 7

References

［1］杨建宇，刘冠军，刘白云，等. 中华中医药道地药材系列汇讲（13）道地药材怀菊花的研究近况［J］. 现代医学与健康研究电子杂志，2020，4（13）：98‑101.
Yang J Y，Liu G J，Liu B Y，et al. A series of lectures on authentic Chinese medicine （13）：recent research on

authentic Chinese medicine Chrysanthemum morifolium［J］. Modern Medicine and Health Research，2020，4（13）：98‑101.

［2］董薇，余永亮，许兰杰，等. 怀菊系列新品种的选育研究［J］. 中药材，2025，48（12）：2976‑2980.

Dong W，Yu Y L，Xu L J，et al. Study on breeding of new varieties of Huaiju series［J］. Journal of Chinese Medicinal Materials，2025，48（12）：2976‑2980.

［3］刘瑞山，谷云. 怀菊花红枣乳酸菌饮料的工艺及稳定性研究［J］. 中国酿造，2013，32（11）：102‑104.

Liu R S，Gu Y. Research on craft and stability of Huai‑Chrysanthemum‑red date Lactobacillus beverage［J］.China Brewing，2013，32（11）：102‑104.

［4］谢玮欢. 农产品电子商务发展策略研究：以焦作市怀菊花为例［J］. 营销界，2022（21）：170‑172.

Xie W H. Study on the development strategy of agricultural products E‑commerce：taking Huaijuhua in Jiaozuo City as an example［J］. Marketing Circles，2022 （21）：170‑172.

［5］徐扬，刘引，彭政，等. 化肥减量配施有机肥对药用菊花产量、品质和药理活性的影响［J］. 应用生态学报，2021，32（8）：2800‑2808.

Xu Y，Liu Y，Peng Z，et al. Effects of chemical fertilizer reduction combined with organic fertilizer on the yield，quality，and pharmacological activity of Chrysanthemum morifolium［J］. Chinese Journal of Applied Ecology，2021，32（8）：2800‑2808.

［6］赫明涛，闫凯旋，尹明明，等. 药用菊花化肥农药使用现状及优化措施［J］. 特种经济动植物，2022，25（1）：119‑120.

He M T，Yan K X，Yin M M，et al. Present situation and optimization measures of chemical fertilizer and pesticide application of medicinal Chrysanthemum［J］.Special Economic Animals and Plants，2022，25（1）：119‑120.

［7］屈凯静，杨钦羽，胡思雨，等. 生物炭在环境污染治理方面的应用研究进展［J］. 北方园艺，2023（16）：123‑130.

Qu K J，Yang Q Y，Hu S Y，et al. Research progress of biochar application in environmental pollution control ［J］. Northern Horticulture，2023（16）：123‑130.

［8］邹雨桐，赵静远，李海霞. 植物源生物炭的制备及其对抗生素的吸附研究进展［J］. 化学与生物工程，2023，40（11）：7‑11.

Zou Y T，Zhao J Y，Li H X. Research progress in preparation and antibiotics adsorption of plant‑derived biochar［J］. Chemistry & Bioengineering，2023，40（11）：7‑11.

［9］柳骁桐. 秸秆生物质炭化还田改土培肥效应研究［D］. 银川：宁夏大学，2020.

Liu X T. Research on biomass carbonization technology［D］. Yinchuan：Ningxia University，2020.

［10］黄兆琴，张乃文，刘霞. 稻壳生物炭的制备及性质表征［J］. 广州化工，2018，46（12）：40‑43.

Huang Z Q，Zhang N W，Liu X. Preparation and characterization of physical and chemical structures of rice husk‑derived biochar［J］. Guangzhou Chemical Industry，2018，46（12）：40‑43.
［11］Chen G，Qiao J，Zhao G H，et al. Rice‑straw biochar regulating effect on Chrysanthemum morifolium Ramat.cv.‘Hangbaiju’［J］. Agronomy Journal，2018，110（5）：1996‑2003.

［12］吴志庄，王道金，厉月桥，等. 施用生物炭肥对黄连木生长及光合特性的影响［J］. 生态环境学报，2015，24（6）：992‑997.

Wu Z Z，Wang D J，Li Y Q，et al. Effects of biochar fertilizer application on growth properties and photosynthetic and physiological characteristics of Pistacia chinensis bunge.［J］. Ecology and Environmental Sciences，2015，24（6）：992‑997.

［13］徐嘉宁，唐姗姗，罗弦，等. 生物炭替代草炭对金盏菊生长和观赏性状的影响［J］. 西北农林科技大学学报（自然科学版），2024，52（10）：119‑126.

Xu J N，Tang S S，Luo X，et al. Effect of biochar substituting peat on growth and ornamental characters of Calendula officinalis［J］. Journal of Northwest A &F University（Natural Science Edition），2024，52（10）：119‑126.

［14］张新俊，杨芳绒，张书文，等. 生物炭对连作土壤性质及菊花生长和品质的影响［J］. 山东农业大学学报（自

然科学版），2022，53（1）：34‑38.

Zhang X J，Yang F R，Zhang S W，et al. Effect of biochar on soil properties of continuous Chrysanthemum morifolium Ramat. cropping and plant growth as well as flower quality［J］. Journal of Shandong Agricultural University（Natural Science Edition），2022，53（1）：34‑38.

［15］张晓波，闫烨. 生物炭施用对菊花连作土壤性质及其生长的影响［J］. 江苏农业科学，2024，52（22）：162‑167.

Zhang X B，Yan Y. Effects of biochar application on soil properties and growth of Chrysanthemum under continuous cropping［J］. Jiangsu Agricultural Sciences，2024，52（22）：162‑167.

［16］武瑞娟，豆叶青. 河南省主要农作物秸秆资源分布及资源化利用需求分析［J］. 可再生能源，2024，42（10）：1289‑1294.

Wu R J，Dou Y Q. Analysis of the distribution and resource utilization needs of main crop straw resources in Henan Province［J］. Renewable Energy Resources，2024，42（10）：1289‑1294.

［17］赵林艳. 不同生物质炭对三七连作障碍的消减效应与机制研究［D］. 昆明：云南师范大学，2022.

Zhao L Y. Effects and mechanisms of biochar derived from different feedstocks to reduce continuous cropping obstacles of Panax notoginseng［D］.Kunming：Yunnan Normal University，2022.

［18］张红娟，陈梦华，何珊珊，等. 有机肥配施生物炭对设施番茄产量、品质及土壤养分和重金属累积的影响［J］. 西北农业学报，2023，32（12）：1978‑1986.

Zhang H J，Chen M H，He S S，et al. Effects of organic fertilizer combined with biochar on yield，quality，soil nutrients and heavy metals in facility tomato［J］. Acta Agriculturae Boreali‑Occidentalis Sinica，2023，32（12）：1978‑1986.

［19］郭平，黄子琪. 河南省农作物秸秆综合利用现状及对策分析［J］. 河南农业，2023（16）：14‑15.
Guo P， Huang Z Q. Present situation and countermeasure analysis of comprehensive utilization of crop straw in Henan Province［J］. Agriculture of Henan，2023（16）：14‑15.

［20］李娇. 河南省粮食作物产业与机械化发展进程研究［J］. 智慧农业导刊，2025，5（4）：48‑51.
Li J. Research on grain crops in Henan Province based on digital agriculture［J］. Journal of Smart Agriculture，2025，5（4）：48‑51.

［21］左开云，阿布力克木·吾斯曼，叶丽努尔·哈德勒，等. 5 个葡萄砧木品种的光合荧光特征分析［J］. 天津农业科学，2023，29（12）：16‑21.
Zuo K Y，Abulkmu W S M，Ylinuer H D L，et al. Analysis on photosynthetic fluorescence characteristics of five grape rootstock varieties［J］. Tianjin Agricultural Sciences，2023，29（12）：16‑21.

［22］吕萌，解静，王跃武，等. 底肥种类和种植密度对“怀菊2号”生长发育、品质及产量的影响［J］. 中药材，2022，
45（4）：771‑777.
Lü M，Xie J，Wang Y W，et al. Effect of base fertilizer type and planting density on growth and development，quality and yield of Chrysanthemum morifolium “Huaiju No. 2”［J］. Journal of Chinese Medicinal Materials，2022，45（4）：771‑777.

［23］鲍士旦. 土壤农化分析［M］. 3 版. 北京：中国农业出版社，2000.
Bao S D. Soil and agricultural chemical analysis ［M］. 3rd ed. Beijing：China Agriculture Press，2000.

［24］王学奎，黄见良. 植物生理生化实验原理与技术［M］. 3 版. 北京：高等教育出版社，2015.
Wang X K，Huang J L. Principles and techniques of plant physiological and biochemical experiments［M］. 3rd ed. Beijing：Higher Education Press，2015.

［25］国家药典委员会. 中华人民共和国药典（一部）［S］. 北京：中国医药科技出版社，2025：322‑323.
National Pharmacopoeia Commission. Pharmacopoeia of the People’s Republic of China（volume Ⅰ）［S］.Beijing：China Pharmaceutical Science and Technology Press，2025：322‑323.

［26］张越. 砂仁中黄酮类化合物的提取及其生物活性研究［D］. 长春：吉林大学，2022.
Zhang Y. Study on extraction and biological activity of flavonoid from Amomum villosum［D］. Changchun：Jilin University，2022.

［27］郭颂，杨卫君，李廷宇，等. 生物炭用量与灌水量对北疆灌区春小麦籽粒产量和品质综合评价［J］. 麦类作物学报，2024，44（5）：658‑666.
Guo S，Yang W J，Li T Y，et al. Comprehensive evaluation of biochar amount and irrigation amount on grain yield and quality of spring wheat in irrigation district of northern Xinjiang［J］. Journal of Triticeae Crops，2024，44（5）：658‑666.

［28］Bharti P，Das A，Roy Choudhury S，et al. Soil aggregation controls biological activities on long‑term integration of organic amendments and inorganic fertilizers in rice‑wheat cropping system［J］.Communications in Soil Science and Plant Analysis，2023，54（13）：1865‑1876.
［29］陈宇琳，吴凤英，张伟婷，等. 不同类型生物炭对芳香樟生长及光合特性的影响［J］. 西北农林科技大学学报（自然科学版），2024，52（2）：70‑80.
Chen Y L，Wu F Y，Zhang W T，et al. Effects of different biochar on growth and photosynthetic characteristics of Cinnamomum camphora［J］. Journal of Northwest A & F University（Natural Science Edition），2024，52（2）：70‑80.

［30］王立业，牛江秀，王真真，等. 景天三七根、茎、花、叶中总黄酮含量测定［J］. 湖北农业科学，2021，60（23）：131‑135.
Wang L Y，Niu J X，Wang Z Z，et al. Determination of total flavonoids in roots，stems，flowers and leaves of Sedum aizoon L.［J］. Hubei Agricultural Sciences，2021，60（23）：131‑135.

［31］戴佩彬. 制备条件对不同原料生物炭理化性质的影响［J］. 生物化工，2021，7（6）：129‑131.
Dai P B. Effect of preparation conditions on physicochemical properties of biochar from different raw materials［J］. Biological Chemical Engineering，2021，7（6）：129‑131.

［32］高凯芳，简敏菲，余厚平，等. 裂解温度对稻秆与稻壳制备生物炭表面官能团的影响［J］. 环境化学，2016，35（8）：1663‑1669.
Gao K F，Jian M F，Yu H P，et al. Effects of pyrolysis temperatures on the biochars and its surface functional groups made from rice straw and rice husk［J］.Environmental Chemistry，2016，35（8）：1663‑1669.

［33］许冬倩. 玉米秸秆生物炭制备及结构特性分析［J］. 广西植物，2018，38（9）：1125‑1135.
Xu D Q. Preparation biochar of corn stover and structural characteristics analysis［J］. Guihaia，2018，38（9）：1125‑1135.

［34］王志丹，代晓华，刘吉利，等. 生物炭对宁夏干旱地区玉米生长及产量的影响［J］. 甘肃农业大学学报，2022，57（1）：74‑82.
Wang Z D，Dai X H，Liu J L，et al. Effect of biochar on growth and yield of maize in arid area of Ningxia［J］.Journal of Gansu Agricultural University，2022，57 （1）：74‑82.

［35］畅芳玲，张沛祯，姚宗路，等. 不同改性玉米秸秆生物炭对黄土丘陵地区土壤水力特性的影响［J］. 环境科学，2024，45（11）：6546‑6554.
Chang F L，Zhang P Z，Yao Z L，et al. Effects of different modified corn straw biochar on soil hydraulic properties in hilly loess areas［J］. Environmental Science，2024，45（11）：6546‑6554.

［36］陈勇. 玉米秸秆还田后土壤质量变化及对作物种子发芽和幼苗生长的影响［D］. 长春：吉林大学，2016.
Chen Y. Changes of soil quality of maize straw after returning to field and its effect on crop seed germination and seedling growth［D］. Changchun：Jilin University，2016.

［37］Wang X B，Song D L，Liang G Q，et al. Maize biochar addition rate influences soil enzyme activity and microbial community composition in a fluvo‑aquic soil［J］. Applied Soil Ecology，2015，96：265‑272.

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