参考文献:

 [1] 岑斌,王慧中.植物蜡质合成与分泌的研究进展［J］.科技通报,2009,25(3):265-271.

［2］Xu X J,Xiao L,Feng J H,et al.Cuticle lipids on heteromorphic leaves of Populus euphratica Oliv.growing in riparian habitats differing in available soil moisture［J］.Physiologia Plantarum,2016,158(3):318-330.

［3］Kunst L,Samuels A L.Biosynthesis and secretion of plant cuticular wax［J］.Progress in Lipid Research,2003,42(1):51-80.

［4］Bernard A,Joubes J.Arabidopsis cuticular waxes:Advances in synthesis,export and regulation［J］.Progress in Lipid Research,2013,52(1):110-129.

［5］Haslam T M,Haslam R,Thoraval D,et al.ECERIFERUM2-LIKE proteins have unique biochemical and physiological functions in verylong-chain fatty acid elongation［J］.Plant Physiology,2015,167(3):682-692.

［6］Rowland O,Zheng H,Hepworth S R,et al.CER4 encodes an alcohol-forming fatty acyl-coenzyme A reductase involved in cuticular wax production in Arabidopsis［J］.Plant Physiology,2006,142(3):866-877.

［7］Bernard A,Domergue F,Pascal S,et al.Reconstitution of plant alkane biosynthesis in yeast demonstrates that Arabidopsis ECERIFERUM1 and ECERIFERUM3 are core components of a very-long-chain alkane synthesis complex［J］.Plant Cell,2012,24(7):3106-3118.

［8］段瑞君,王爱东,陈国雄.植物角质层基因研究进展［J］.植物学报,2017,52(5):637-651.

［9］Panikashvili D,Shi J X,Schreiber L,et al.The Arabidopsis ABCG13 transporter is required for flower cuticle secretion and patterning of the petal epidermis［J］.New Phytologist,2011,190(1):113-124.

［10］胡晓敏,张志飞,饶力群,等.植物角质层蜡质合成与调控的分子生物学研究进展［J］.武汉植物学研究,2007,25(4):377-380.

［11］Kim H,Lee S B,Kim H J,et al.Characterization of glycosylphosphatidylinositol-anchored lipid transfer protein 2(LTPG2)and overlapping function between LTPG/LTPG1 and LTPG2 in cuticular wax export or accumulation in Arabidopsis thaliana［J］.Plant and Cell Physiology,2012,53(8):1391-1403.

［12］Lee S B,Suh M C.Cuticular wax biosynthesis is up-regulated by the MYB94 transcription factor in Arabidopsis［J］.Plant and Cell Physiology,2015,56(1):48-60.

［13］Liang G,He H,Li Y,et al.MYB82 functions in regulation of trichome development in Arabidopsis［J］.Journal of Experimental Botany,2014,65(12):3215-3223.

［14］许发喜,刘翠芳,邹杰,等.植物角质层对非生物逆境胁迫响应研究进展［J］.中国生物工程杂志,2010,30(8):126-130.

［15］邓彦斌,姜彦成,刘健.新疆10种藜科植物叶片和同化枝的旱生和盐生结构的研究［J］.植物生态学报,1998,22(2):164-170.

［16］Shepherd T,Wynne G D.The effects of stress on plant cuticular waxes［J］.New Phytol,2006,171(3):469-499.

［17］Ladaniya M S.Physicochemical,respiratory and fungicide residue changes in wax coated mandarin fruit stored at chilling temperature with intermittent warming［J］.Journal of Food Science and Technology-Mysore,2011,48(2):150-158.

［18］Dou H.Effect of coating application on chilling injury of grapefruit cultivars［J］.Horticulture Science,2004,39(3):558-561.

［19］Taji T,Seki M,Satou M,et al.Comparative genomics in salt tolerance between Arabidopsis and Arabidopsis-related halophyte salt cress using Arabidopsis microarray［J］.Plant Physiology,2004,135(3):1697-1709.

［20］Griffith M,Timonin M,Wong A C,et al.Thellungiella:An Arabidopsis-related model plant adapted to cold temperatures［J］.Plant Cell and Environment,2007,30(5):529-538.

［21］Rahman L N,Bamm V V,Voyer J A,et al.Zinc induces disorder-to-order transitions in free and membrane-associated Thellungiella salsuginea dehydrins TsDHN-1 and TsDHN-2:A solution CD and solid-state ATR-FTIR study［J］.Amino Acids,2011,40(5):1485-1502.

［22］Li P H,Mane S P,Sioson A A,et al.Effects of chronic ozone exposure on gene expression in Arabidopsis thaliana ecotypes and in Thellungielia halophila［J］.Plant Cell and Environment,2006,29(5):854-868.

［23］Kant S,Bi YM,Weretilnyk E,et al.The Arabidopsis halophytic relative Thellungiella halophila tolerates nitrogen-limiting conditions by maintaining growth,nitrogen uptake,and assimilation［J］.Plant Physiology,2008,147(3):1168-1180.

［24］赵宝.三种生态型盐芥表皮蜡质及其合成相关的KCS家族各基因表达［D］.济南:山东师范大学,2007.

［25］Zhang X D,Wang R P,Zhang F J,et al.Lipid profiling and tolerance to low-temperature stress in Thellungiella salsuginea in comparison with Arabidopsis thaliana［J］.Biologia Plantarum,2013,57(1):149-153.

［26］王增兰.盐芥耐盐生理及分子生物学基础研究［D］.济南:山东师范大学,2004.

［27］Inan G,Zhang Q,Li P,et al.Salt cress.A halophyte and cryophyte Arabidopsis relative model system and its applicability to molecular genetic analyses of growth and development of extremophiles［J］.Plant Physiology,2004,135(3):1718-1737.

［28］Mittler R.Oxidative stress,antioxidants and stress tolerance［J］.Trends in Plant Science,2002,7(9):405-410.

［29］倪郁,宋超,王小清.低温胁迫下拟南芥表皮蜡质的响应机制［J］.中国农业科学,2014,47(2):252-261.

［30］Chen X,Goodwin S M,Boroff V L,et al.Cloning and characterization of the WAX2 gene of Arabidopsis involved in cuticle membrane and wax production［J］.Plant Cell,2003,15(5):1170-1185.

［31］Livak K J,Schmittgen T D.Analysis of relative gene expression data using real-time quantitative PCR and the 2-△△CT method［J］.Methods,2001,25(4):402-408.

［32］Kosma D K,Bourdenx B,Bernard A,et al.The impact of water deficiency on leaf cuticle lipids of Arabidopsis［J］.Plant Physiology,2009,151(4):1918-1929.

［33］Aharoni A,Dixit S,Jetter R,et al.The SHINE clade of AP2 domain transcription factors activates wax biosynthesis,alters cuticle properties,and confers drought tolerance when overexpressed in Arabidopsis［J］.Plant Cell,2004,16(9):2463-2480.

［34］Bourdenx B,Bernard A,Domergue F,et al.Overexpression of Arabidopsis ECERIFERUM1 promotes wax very-long-chain alkane biosynthesis and influences plant response to biotic and abiotic stresses［J］.Plant Physiology,2011,156(1):29-45.

［35］Seo P J,Lee S B,Suh M C,et al.The MYB96 transcription factor regulates cuticular wax biosynthesis under drought conditions in Arabidopsis［J］.Plant Cell,2011,23(3):1138-1152.

［36］Jenks M A,Andersen L,Teusink R S,et al.Leaf cuticular waxes of potted rose cultivars as affected by plant development,drought and paclobutrazol treatments［J］.Physiol Plant,2001,112(1):62-70.

［37］Cameron K D,Teece M A,Smart L B.Increased accumulation of cuticular wax and expression of lipid transfer protein in response to periodic drying events in leaves of tree tobacco［J］.Plant Physiology,2006,140(1):176-183.

［38］Kim K S,Park S H,Jenks M A.Changes in leaf cuticular waxes of sesame(Sesamum indicum L.)plants exposed to water deficit［J］.Journal of Plant Physiology,2007,164(9):1134-1143.

［39］Kim K,Park S,Jenks M,et al.Influence of water deficit on leaf cuticular waxes of soybean［J］.International Journal of Plant Sciences,2006,51(3):307-316.

［40］Borisjuk N,Hrmova M,Lopato S.Transcriptional regulation of cuticle biosynthesis［J］.Biotechnology Advances,2014,32(2):526-540.

［41］Le Provost G,Domergue F,Lalanne C,et al.Soil water stress affects both cuticular wax content and cuticle-related gene expression in young saplings of maritime pine(Pinus pinaster Ait)［J］.BMC Plant Biology,2013,13(1):95-107.

［42］McFarlane H E,Shin J J,Bird D A,et al.Arabidopsis ABCG transporters,which are required for export of diverse cuticular lipids,dimerize in different combinations［J］.Plant Cell,2010,22(9):3066-3075.

［43］Shinozaki K,Yamaguchi-Shinozaki K,Seki M.Regulatory network of gene expression in the drought and cold stress responses［J］.Current Opinion in Plant Biology,2003,6(5):410-417.

［44］Zhu J K.Salt and drought stress signal transduction in plants［J］.Annual Review of Plant Biology,2002,53(53):247-273.

［45］Lee S B,Kim H U,Suh M C.MYB94 and MYB96 additively activate cuticular wax biosynthesis in Arabidopsis［J］.Plant Cell Physiology,2016,57(11):2300-2311.

［46］Lu S,Zhao H,Des Marais D L,et al.Arabidopsis ECERIFERUM9 involvement in cuticle formation and maintenance of plant water status［J］.Plant Physiology,2012,159(3):930-944.

［47］Xu X J,Feng J C,Lu S Y,et al.Leaf cuticular lipids on the Shandong and Yukon ecotypes of saltwater cress,Eutrema salsugineum,and their response to water deficiency and impact on cuticle permeability［J］.Physiologia Plantarum,2014,151(4):446-458.